Raymond D. Semlitsch, David E. Scott, Joseph H. K. Pechmann, J. Whitfield Gibbons, Phenotypic Variation in the Arrival Time of Breeding Salamanders: Individual Repeatability and Environmental Influences, Journal of Animal Ecology, Vol. 62, No. 2 (Apr., 1993), pp. 334-340
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Semlitsch et al. (1993) studied this question.
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