Timothy M. Goater, Gerald W. Esch, Albert O. Bush, Helminth Parasites of Sympatric Salamanders: Ecological Concepts at Infracommunity, Component and Compound Community Levels, The American Midland Naturalist, Vol. 118, No. 2 (Oct., 1987), pp. 289-300
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Goater et al. (1987) studied this question.
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