Yi Tao, Fenghua Yuan, R. Todd Leister, Frederick M. Ausubel, Fumiaki Katagiri, Mutational Analysis of the Arabidopsis Nucleotide Binding Site: Leucine-Rich Repeat Resistance Gene RPS2, The Plant Cell, Vol. 12, No. 12 (Dec., 2000), pp. 2541-2554
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Yi et al. (2000) studied this question.