W. D. Stock, J. S. Pate, J. Delfs, Influence of Seed Size and Quality on Seedling Development Under Low Nutrient Conditions in Five Australian and South African Members of the Proteaceae, Journal of Ecology, Vol. 78, No. 4 (Dec., 1990), pp. 1005-1020
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Stock et al. (1990) studied this question.
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