Lennart Persson, Lars Johansson, Gunnar Andersson, Sebastian Diehl, Stellan F. Hamrin, Density Dependent Interactions in Lake Ecosystems: Whole Lake Perturbation Experiments, Oikos, Vol. 66, No. 2 (Mar., 1993), pp. 193-208
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Persson et al. (1993) studied this question.