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Ecological energetics can be a useful approach with which to assess the importance of different species populations in their contribution to the structure, productivity and functioning of communities (Macfadyen 1963; Odum 1963; Slobodkin 1962). Theoretical considerations of ecological energetics have been treated fully by Lindemann (1942), Phillipson (1966), Odum (1963) and Slobodkin (1962); however, the paucity of field data emphasizes the need for more research (Engelmann 1966). Following I.B.P. terminology (Petrusewicz 1967; Ricker 1968), the energy budget of a population may be summarized by the equation:
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Roger N. Hughes (1970) studied this question.
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