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I. R. Wallis, B. T. Henen, K. A. Nagy, Egg Size and Annual Egg Production by Female Desert Tortoises (Gopherus agassizii): The Importance of Food Abundance, Body Size, and Date of Egg Shelling, Journal of Herpetology, Vol. 33, No. 3 (Sep., 1999), pp. 394-408
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Wallis et al. (1999) studied this question.
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