Ann M. McLuckie, Trip Lamb, Cecil R. Schwalbe, Robert D. McCord, Genetic and Morphometric Assessment of an Unusual Tortoise (Gopherus agassizii) Population in the Black Mountains of Arizona, Journal of Herpetology, Vol. 33, No. 1 (Mar., 1999), pp. 36-44
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McLuckie et al. (1999) studied this question.
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