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Kei'ichiroh Iguchi, Motoyoshi Yamaguchi, Adaptive Significance of Inter- and Intrapopulational Egg Size Variation in Ayu Plecoglossus altivelis (Osmeridae), Copeia, Vol. 1994, No. 1 (Feb. 1, 1994), pp. 184-190
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Iguchi et al. (1994) studied this question.
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