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Barry J. F. Biggs, Nancy C. Tuchman, Rex L. Lowe, R. Jan Stevenson, Resource Stress Alters Hydrological Disturbance Effects in a Stream periphyton Community, Oikos, Vol. 85, No. 1 (Apr., 1999), pp. 95-108
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Biggs et al. (1999) studied this question.
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