G. R. Coope, R. B. Angus, An Ecological Study of a Temperate Interlude in the Middle of the Last Glaciation, Based on Fossil Coleoptera from Isleworth, Middlesex, Journal of Animal Ecology, Vol. 44, No. 2 (Jun., 1975), pp. 365-391
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Coope et al. (1975) studied this question.
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