D. G. Bedo, Cytogenetics and Evolution of Simulium ornatipes Skuse (Diptera: Simuliidae). II. Temporal Variation in Chromosomal Polymorphisms and Homosequential Sibling Species, Evolution, Vol. 33, No. 1, Part 2 (Mar., 1979), pp. 296-308
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D. G. Bedo (1979) studied this question.
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