The primary productivity of the benthic microflora was measured using in situ techniques. Primary productivity ranged from below 0.5 mg C m−2 h−1 in winter when the sampling area was covered with ice to nearly 57.0 mg C m−2 h−1 in August. This is 8 times the productivity of the ice algae and twice that of the phytoplankton.
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Matheke et al. (1974) studied this question.