Robert G. Gregerson, Susan S. Miller, Scott N. Twary, J. Stephen Gantt, Carroll P. Vance, Molecular Characterization of NADH-Dependent Glutamate Synthase from Alfalfa Nodules, The Plant Cell, Vol. 5, No. 2 (Feb., 1993), pp. 215-226
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Gregerson et al. (1993) studied this question.