Kenneth D. Johnson, , Maarten J. Chrispeels, An Intrinsic Tonoplast Protein of Protein Storage Vacuoles in Seeds Is Structurally Related to a Bacterial Solute Transporter (GlpF), The Plant Cell, Vol. 2, No. 6 (Jun., 1990), pp. 525-532
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Johnson et al. (1990) studied this question.