Gregory B. Wilson, Theodore L. Jahn, James R. Fonseca, Studies on Ciliary Beating of Frog Pharyngeal Epithelium in vitro: I. Isolation and Ciliary Beat of Single Cells, Transactions of the American Microscopical Society, Vol. 94, No. 1 (Jan., 1975), pp. 43-57
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Wilson et al. (1975) studied this question.
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