Up to the present, light microscopy has shown the middle lamella between plant cells as an amorphous, intercellular substance (Esau, 1965). Furthermore, this concept of middle-lamellar structure has been strengthened by the few available micrographs from electron microscopy. The micrographs seem to have been chosen to emphasize cell wall microfibrils rather than to show the components of the surrounding matrix. Hence, an
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Leppard et al. (1971) studied this question.
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