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I analyzed formations of Canada Geese (Branta canadensis) with a single, direct method of testing predictions from multiple hypotheses. The results support both energetic (aerodynamic) advantage and orientation communication through visual contact as functions of this complex behavior. Comparison of observed positioning patterns with criteria for optimal function suggests priority may be given to the maximization of energy savings within limits imposed by environmental and other constraints.
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John P. Badgerow (1988) studied this question.
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