OUGHEST is a sex-linked, recessive character of Drosophila melano-R gaster manifested phenotypically in bulging, roughened eyes that have facets irregular in size and shape. The allele roughest3 (mi3) was obtained by GRUNEBERG (1935) from an X-ray experiment. Eyes of rst3 flies show mosaics of smooth and rough facets with rough patches more pronounced in the males than in the females. Roughness is accentuated if flies are raised at temperatures between 16 and 19C; at higher temperatures the females more frequently overlap the wild type in appearance. Viability and fertility are good in both sexes. Genetic analysis (GRUNE-BERG 1935) indicated that rst3 is associated with a long inversion in the GENETICS 27: 537 Sept.
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Berwind P. Kaufmann (1942) studied this question.