A.-C. Hansson, R. Pettersson, Uptake and Above- and Below-Ground Allocation of Soil Mineral-N and Fertilizer-15N in a Perennial Grass Ley (Festuca pratensis), Journal of Applied Ecology, Vol. 26, No. 1 (Apr., 1989), pp. 259-271
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Hansson et al. (1989) studied this question.