The Nerthra alaticollis species-group is defined as a monophyletic group on the basis of wing and genitalic characters. Two new species are described: Nerthra appha n. sp. and N. polhemi n.sp. Nerthra membranacea Nieser is recognized as a junior synonym of N. adspersa. Nerthra tuberculata (Montandon) and N. praecipua Todd are removed from the alaticollis species-group. Nine species are assigned to the species-group, and all species are described with illustrations. A phylogeny of species is provided.
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Cassis et al. (2001) studied this question.
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