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N. O. Bianchi, O. A. Reig, O. J. Molina, F. N. Dulout, Cytogenetics of the South American Akodont Rodents (Cricetidae). I. A Progress Report of Argentinian and Venezuelan Forms, Evolution, Vol. 25, No. 4 (Dec., 1971), pp. 724-736
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Bianchi et al. (1971) studied this question.
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