W. W. Marquardt, S. K. Kadavil, D. B. Snyder, Comparison of Ciliary Activity and Virus Recovery from Tracheas of Chickens and Humoral Immunity after Inoculation with Serotypes of Avian Infectious Bronchitis Virus, Avian Diseases, Vol. 26, No. 4 (Oct. - Dec., 1982), pp. 828-834
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Marquardt et al. (1982) studied this question.