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P. V. Biscoe, J. N. Gallagher, E. J. Littleton, J. L. Monteith, R. K. Scott, Barley and its Environment. IV. Sources of Assimilate for the Grain, Journal of Applied Ecology, Vol. 12, No. 1 (Apr., 1975), pp. 295-318
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Biscoe et al. (1975) studied this question.
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