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Raymond D. Semlitsch, , Gaston-Denis Guex, Competition Among Tadpoles of Coexisting Hemiclones of Hybridogenetic Rana esculenta: Support for the Frozen Niche Variation Model, Evolution, Vol. 51, No. 4 (Aug., 1997), pp. 1249-1261
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Semlitsch et al. (1997) studied this question.
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