S. J. Milton, W. R. Siegfried, W. R. J. Dean, The Distribution of Epizoochoric Plant Species: A Clue to the Prehistoric Use of Arid Karoo Rangelands by Large Herbivores, Journal of Biogeography, Vol. 17, No. 1 (Jan., 1990), pp. 25-34
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Milton et al. (1990) studied this question.
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