J. Silvertown, M. Tremlett, Interactive Effects of Disturbance and Shade upon Colonization of Grassland: An Experiment with Anthriscus sylvestris (L.) Hoffm., Conium maculatum L., Daucus carota L. and Heracleum sphondylium L., Functional Ecology, Vol. 3, No. 2 (1989), pp. 229-235
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Silvertown et al. (1989) studied this question.
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