Alvan A. Karlin, Kathy S. Williams, Kimberly G. Smith, Derrick W. Sugg, Biochemical Evidence for Rapid Changes in Heterozygosity in a Population of Periodical Cicadas (Magicicada tredecassini), The American Midland Naturalist, Vol. 125, No. 2 (Apr., 1991), pp. 213-221
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Karlin et al. (1991) studied this question.
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