Denis H. Lynn, Jacques Berger, The Thyrophylacidae, a Family of Carnivorous Philasterine Ciliates Entocommensal in Strongylocentrotid Echinoids, Transactions of the American Microscopical Society, Vol. 92, No. 4 (Oct., 1973), pp. 533-557
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Lynn et al. (1973) studied this question.
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