Flounders captured in summer in the intertidal zone at high tide fed mainly on food species that live in that zone. Flounders captured in the subtidal zone fed partially on food species that live there, and partially on intertidal food species. From these feeding habits, and from the magnitude of the movement into the intertidal zone (previously reported), we conclude that large intertidal areas are major feeding grounds for winter flounder. The quantities of filamentous algae in the stomachs suggest that winter flounders can utilize plant material.
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Wells et al. (1973) studied this question.