Nils Ryman, Ramone Baccus, Christina Reuterwall, Michael H. Smith, Effective Population Size, Generation Interval, and Potential Loss of Genetic Variability in Game Species under Different Hunting Regimes, Oikos, Vol. 36, No. 3 (Mar., 1981), pp. 257-266
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Ryman et al. (1981) studied this question.
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