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The simultaneous formation of E and Z double bonds results from the syn elimination of H and/or D atoms from different conformations of 4-hydroxysphinganine [D4]1. Δ8-Sphingolipid desaturase from Helianthus annuus is heterologously expressed in yeast and catalyzes the transformation to E olefin 2 (88%) and Z olefin 3 (12%).
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Beckmann et al. (2002) studied this question.
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