Rosemary Carpenter, Lucy Copsey, Coral Vincent, Sandra Doyle, Ruth Magrath, Enrico Coen, Control of Flower Development and Phyllotaxy by Meristem Identity Genes in Antirrhinum, The Plant Cell, Vol. 7, No. 12 (Dec., 1995), pp. 2001-2011
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Carpenter et al. (1995) studied this question.