Elizabeth A. Kellogg, Rudi Appels, Roberta J. Mason-Gamer, When Genes Tell Different Stories: The Diploid Genera of Triticeae (Gramineae), Systematic Botany, Vol. 21, No. 3 (Jul. - Sep., 1996), pp. 321-347
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Kellogg et al. (1996) studied this question.
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