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Charles Kwit, Mark W. Schwartz, William J. Platt, James P. Geaghan, The Distribution of Tree Species in Steepheads of the Apalachicola River Bluffs, Florida, The Journal of the Torrey Botanical Society, Vol. 125, No. 4 (Oct. - Dec., 1998), pp. 309-318
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Kwit et al. (1998) studied this question.
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