Robert K. Kuzoff, Douglas E. Soltis, Larry Hufford, Pamela S. Soltis, Phylogenetic Relationships Within Lithophragma (Saxifragaceae): Hybridization, Allopolyploidy, and Ovary Diversification, Systematic Botany, Vol. 24, No. 4 (Oct. - Dec., 1999), pp. 598-615
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Kuzoff et al. (1999) studied this question.
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