Hexagyne philippiana gen. et sp. nov., a piperalean angiosperm from the Early Cretaceous of northern Gondwana (Crato Formation, Brazil)

Hexagyne philippiana gen. et sp. nov. represents the first macrofossil piperalean taxon from the Early Cretaceous (Aptian). The fossil material from northern Gondwana consists of a twig with well‐preserved leaves and a characteristic venation pattern plus a twig with attached leaves and several flowers combined in a raceme. The new taxon shares many characters, such as leaf shape and venation pattern, as well as flower morphology, including a clearly developed perianth and six fused carpels, with extant Aristolochiaceae (sensu lato) and/or related clades. Representatives of Aristolochiaceae grow today in Laurasia. The Early Cretaceous finds in South America (Brazil) may indicate a Gondwanan origin of this group. Based on its morphology, Hexagyne gen. nov. might have had an ecology of an understory plant in a shaded environment.