Primary production of benthic microalgae was studied in an intertidal sand flat in the Ria de Arosa (NW Spain), where a n intensive bivalve aquaculture takes place. Active and degraded pigments, organic matter, and vertical migration of algae were also studied. Total annual production. combining high and low tide values, reached 79 g C m-2, while water column production in the same place was only 6 gC m-2, which suggests that resuspended benthic microalgae can be a major food source for bivalves. Year-round production showed no clear seasonal peaks, probably due to intense sediment d~sturbance by culturing manipulations, and to high detrital supply: 94 to 98 % of organic matter in the sediment is allochtonous non-living material.
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Varela et al. (1985) studied this question.
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