An investigation of the dune-living spiders was made during 1960-1962 in a coastal sand dune area of 40 ha at Sandhammaren, Province SkAne, Sweden. Eleven subdivisions were recognized for a vegetational survey; 821 samples from areas of 1 m2 each collected. 33 875 spiders were obtained, belonging to 152 species of which 80 were accidental species. Testing by means of an association method based on presence or absence data for 35 species in all samples, seven association groups were distinguished. By means of another grouping method using distance functions with an adjustable index, taxicoenoses, and their habitats were fixed.
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Sven Almquist (1973) studied this question.