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David C. Heins, Matthew D. Machado, Spawning Season, Clutch Characteristics, Sexual Dimorphism and Sex Ratio in the Redfin Darter Etheostoma whipplei, The American Midland Naturalist, Vol. 129, No. 1 (Jan., 1993), pp. 161-171
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Heins et al. (1993) studied this question.
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