Chikako Sato, Kiyohide Kojima, Taiju Matsuzawa, Yorio Hinuma, Relationship between Loss of Negative Charge on Nuclear Membrane and Loss of Colony-Forming Ability in X-Irradiated Cells, Radiation Research, Vol. 62, No. 2 (May, 1975), pp. 250-257
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Sato et al. (1975) studied this question.