Paul A. Opler, Gordon W. Frankie, Herbert G. Baker, Comparative Phenological Studies of Treelet and Shrub Species in Tropical Wet and Dry Forests in the Lowlands of Costa Rica, Journal of Ecology, Vol. 68, No. 1 (Mar., 1980), pp. 167-188
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Opler et al. (1980) studied this question.
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