Zora Modrusan, Leonore Reiser, Kenneth A. Feldmann, Robert L. Fischer, George W. Haughn, Homeotic Transformation of Ovules into Carpel-Like Structures in Arabidopsis, The Plant Cell, Vol. 6, No. 3 (Mar., 1994), pp. 333-349
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Modrušan et al. (1994) studied this question.