Raymond D. Semlitsch, Gary B. Moran, Ecology of the Redbelly Snake (Storeria occipitomaculata) Using Mesic Habitats in South Carolina, The American Midland Naturalist, Vol. 111, No. 1 (Jan., 1984), pp. 33-40
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Semlitsch et al. (1984) studied this question.
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