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L. D. Gottlieb, V. S. Ford, Phylogenetic Relationships Among the Sections of Clarkia (Onagraceae) Inferred from the Nucleotide Sequences of PgiC, Systematic Botany, Vol. 21, No. 1 (Jan. - Mar., 1996), pp. 45-62
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Gottlieb et al. (1996) studied this question.
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