Ingrid M. van der Meer, Maike E. Stam, Arjen J. van Tunen, Joseph N. M. Mol, Antoine R. Stuitje, Antisense Inhibition of Flavonoid Biosynthesis in Petunia Anthers Results in Male Sterility, The Plant Cell, Vol. 4, No. 3 (Mar., 1992), pp. 253-262
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Meer et al. (1992) studied this question.