Claude W. dePamphilis, Robert Wyatt, Hybridization and Introgression in Buckeyes (Aesculus: Hippocastanaceae): A Review of the Evidence and a Hypothesis to Explain Long-Distance Gene Flow, Systematic Botany, Vol. 14, No. 4 (Oct. - Dec., 1989), pp. 593-611
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dePamphilis et al. (1989) studied this question.
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