Thomas Uzzell, Ilya S. Darevsky, Biochemical Evidence for the Hybrid Origin of the Parthenogenetic Species of the Lacerta saxicola Complex (Sauria: Lacertidae), with a Discussion of Some Ecological and Evolutionary Implications, Copeia, Vol. 1975, No. 2 (May 23, 1975), pp. 204-222
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Uzzell et al. (1975) studied this question.
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