W. B. Watt, C. Kremen, P. Carter, Testing the `Mimicry' Explanation for the Colias `alba' Polymorphism: Patterns of Co-Occurrence of Colias and Pierine Butterflies, Functional Ecology, Vol. 3, No. 2 (1989), pp. 193-199
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Watt et al. (1989) studied this question.
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