Hugh C. Cutler, Marian C. Cutler, Studies on the Structure of the Maize Plant, Annals of the Missouri Botanical Garden, Vol. 35, No. 4, Special Number: The 1947 Conference on the Inflorescences of Zea Mays (Nov., 1948), pp. 301-316
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Cutler et al. (1948) studied this question.
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